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TERRESTRIAL, COASTAL AND FRESHWATER ECOSYSTEMS – HABITAT TYPES AND GROUPS OF HABITAT TYPES REFERRED TO IN ARTICLE 4(1) AND (4) The list below includes all terrestrial, coastal and freshwater habitat types listed in Annex I to Directive 92/43/EEC referred to in Article 4(1) and (4), as well as six groups of those habitat types, namely 1) wetlands (coastal and inland), 2) grasslands and other pastoral habitats, 3) river, lake, alluvial and riparian habitats, 4) forests, 5) steppe, heath and scrub habitats and 6) rocky and dune habitats.

1. GROUP 1: Wetlands (coastal & inland)

Habitat type code as referred to in Annex I to Directive 92/43/EEC Habitat type name as referred to in Annex I to Directive 92/43/EEC Coastal and salt habitats 1130 Estuaries 1140 Mudflats and sandflats not covered by seawater at low tide 1150 Coastal lagoons 1310 Salicornia and other annuals colonizing mud and sand 1320 Spartina swards (Spartinion maritimae) 1330 Atlantic salt meadows (Glauco-Puccinellietalia maritimae) 1340 Inland salt meadows 1410 Mediterranean salt meadows (Juncetalia maritimi) 1420 Mediterranean and thermo-Atlantic halophilous scrubs (Sarcocornetea fruticosi) 1530 Pannonic salt steppes and salt marshes 1650 Boreal Baltic narrow inlets Wet heaths and peat grassland 4010 Northern Atlantic wet heaths with Erica tetralix 4020 Temperate Atlantic wet heaths with Erica ciliaris and Erica tetralix 6460 Peat grasslands of Troodos Mires, bogs and fens 7110 Active raised bogs 7120 Degraded raised bogs still capable of natural regeneration 7130 Blanket bogs 7140 Transition mires and quaking bogs 7150 Depressions on peat substrates of the Rhynchosporion 7160 Fennoscandian mineral-rich springs and springfens 7210 Calcareous fens with Cladium mariscus and species of the Caricion davallianae 7220 Petrifying springs with tufa formation (Cratoneurion) 7230 Alkaline fens 7240 Alpine pioneer formations of the Caricion bicoloris-atrofuscae 7310 Aapa mires 7320 Palsa mires Wet forests 9080 Fennoscandian deciduous swamp woods 91D0 Bog woodland

2. GROUP 2: Grasslands and other pastoral habitats

Habitat type code as referred to in Annex I to Directive 92/43/EEC Habitat type name as referred to in Annex I to Directive 92/43/EEC Costal and dune habitats 1630 Boreal Baltic coastal meadows 21A0 Machairs Heath and scrub habitats 4030 European dry heaths 4040 Dry Atlantic coastal heaths with Erica vagans 4090 Endemic oro-Mediterranean heaths with gorse 5130 Juniperus communis formations on heaths or calcareous grasslands 8240 Limestone pavements Grasslands 6110 Rupicolous calcareous or basophilic grasslands of the Alysso-Sedion albi 6120 Xeric sand calcareous grasslands 6130 Calaminarian grasslands of the Violetalia calaminariae 6140 Siliceous Pyrenean Festuca eskia grasslands 6150 Siliceous alpine and boreal grasslands 6160 Oro-Iberian Festuca indigesta grasslands 6170 Alpine and subalpine calcareous grasslands 6180 Macaronesian mesophile grasslands 6190 Rupicolous pannonic grasslands (Stipo-Festucetalia pallentis) 6210 Semi-natural dry grasslands and scrubland facies on calcareous substrates (Festuco-Brometalia) 6220 Pseudo-steppe with grasses and annuals of the Thero-Brachypodietea 6230 Species-rich Nardus grasslands, on silicious substrates in mountain areas (and submountain areas in Continental Europe) 6240 Sub-Pannonic steppic grasslands 6250 Pannonic loess steppic grasslands 6260 Pannonic sand steppes 6270 Fennoscandian lowland species-rich dry to mesic grasslands 6280 Nordic alvar and precambrian calcareous flatrocks 62A0 Eastern sub-Mediterranean dry grasslands (Scorzoneratalia villosae) 62B0 Serpentinophilous grassland of Cyprus 62C0 Ponto-Sarmatic steppes 62D0 Oro-Moesian acidophilous grasslands 6410 Molinia meadows on calcareous, peaty or clayey-silt-laden soils (Molinion caeruleae) 6420 Mediterranean tall humid grasslands of the Molinio-Holoschoenion 6510 Lowland hay meadows (Alopecurus pratensis, Sanguisorba officinalis) 6520 Mountain hay meadows Dehesas and wooded meadows 6310 Dehesas with evergreen Quercus spp

6530 Fennoscandian wooded meadows 9070 Fennoscandian wooded pastures 3. GROUP 3: River, lake, alluvial and riparian habitats Habitat type code as referred to in Annex I to Directive 92/43/EEC Habitat type name as referred to in Annex I to Directive 92/43/EEC Rivers and lakes 3110 Oligotrophic waters containing very few minerals of sandy plains (Littorelletalia uniflorae) 3120 Oligotrophic waters containing very few minerals generally on sandy soils of the West Mediterranean, with Isoetes spp 3130 Oligotrophic to mesotrophic standing waters with vegetation of the Littorelletea uniflorae and/or of the Isoëto-Nanojuncetea 3140 Hard oligo-mesotrophic waters with benthic vegetation of Chara spp

3150 Natural eutrophic lakes with Magnopotamion or Hydrocharition — type vegetation 3160 Natural dystrophic lakes and ponds 3170 Mediterranean temporary ponds 3180 Turloughs 3190 Lakes of gypsum karst 31A0 Transylvanian hot-spring lotus beds 3210 Fennoscandian natural rivers 3220 Alpine rivers and the herbaceous vegetation along their banks 3230 Alpine rivers and their ligneous vegetation with Myricaria germanica 3240 Alpine rivers and their ligneous vegetation with Salix elaeagnos 3250 Constantly flowing Mediterranean rivers with Glaucium flavum 3260 Water courses of plain to montane levels with the Ranunculion fluitantis and Callitricho-Batrachion vegetation 3270 Rivers with muddy banks with Chenopodion rubri p.p. and Bidention p.p. vegetation 3280 Constantly flowing Mediterranean rivers with Paspalo-Agrostidion species and hanging curtains of Salix and Populus alba 3290 Intermittently flowing Mediterranean rivers of the Paspalo-Agrostidion 32A0 Tufa cascades of karstic rivers of the Dinaric Alps Alluvial meadows 6430 Hydrophilous tall herb fringe communities of plains and of the montane to alpine levels 6440 Alluvial meadows of river valleys of the Cnidion dubii 6450 Northern boreal alluvial meadows 6540 Sub-Mediterranean grasslands of the Molinio-Hordeion secalini Alluvial/Riparian forests 9160 Sub-Atlantic and medio-European oak or oak-hornbeam forests of the Carpinion betuli 91E0 Alluvial forests with Alnus glutinosa and Fraxinus excelsior (Alno-Padion, Alnion incanae, Salicion albae) 91F0 Riparian mixed forests of Quercus robur, Ulmus laevis and Ulmus minor, Fraxinus excelsior or Fraxinus angustifolia, along the great rivers (Ulmenion minoris) 92A0 Salix alba and Populus alba galleries 92B0 Riparian formations on intermittent Mediterranean water courses with Rhododendron ponticum, Salix and others 92C0 Platanus orientalis and Liquidambar orientalis woods (Platanion orientalis) 92D0 Southern riparian galleries and thickets (Nerio-Tamaricetea and Securinegion tinctoriae) 9370 Palm groves of Phoenix

Habitat type code as referred to in Annex I to Directive 92/43/EEC Habitat type name as referred to in Annex I to Directive 92/43/EEC Boreal forests 9010 Western Taïga 9020 Fennoscandian hemiboreal natural old broad-leaved deciduous forests (Quercus, Tilia, Acer, Fraxinus or Ulmus) rich in epiphytes 9030 Natural forests of primary succession stages of landupheaval coast 9040 Nordic subalpine/subarctic forests with Betula pubescens ssp. czerepanovii 9050 Fennoscandian herb-rich forests with Picea abies 9060 Coniferous forests on, or connected to, glaciofluvial eskers Temperate forests 9110 Luzulo-Fagetum beech forests 9120 Atlantic acidophilous beech forests with Ilex and sometimes also Taxus in the shrublayer (Quercion robori-petraeae or Ilici-Fagenion) 9130 Asperulo-Fagetum beech forests 9140 Medio-European subalpine beech woods with Acer and Rumex arifolius 9150 Medio-European limestone beech forests of the Cephalanthero-Fagion 9170 Galio-Carpinetum oak-hornbeam forests 9180 Tilio-Acerion forests of slopes, screes and ravines 9190 Old acidophilous oak woods with Quercus robur on sandy plains 91A0 Old sessile oak woods with Ilex and Blechnum in the British Isles 91B0 Thermophilous Fraxinus angustifolia woods 91G0 Pannonic woods with Quercus petraea and Carpinus betulus 91H0 Pannonian woods with Quercus pubescens 91I0 Euro-Siberian steppic woods with Quercus spp. 91J0 Taxus baccata woods of the British Isles 91K0 Illyrian Fagus sylvatica forests (Aremonio-Fagion) 91L0 Illyrian oak-hornbeam forests (Erythronio-Carpinion) 91M0 Pannonian-Balkanic turkey oak – sessile oak forests 91P0 Holy Cross fir forest (Abietetum polonicum) 91Q0 Western Carpathian calcicolous Pinus sylvestris forests 91R0 Dinaric dolomite Scots pine forests (Genisto januensis-Pinetum) 91S0 Western Pontic beech forests 91T0 Central European lichen Scots pine forests 91U0 Sarmatic steppe pine forest 91V0 Dacian Beech forests (Symphyto-Fagion) 91W0 Moesian beech forests 91X0 Dobrogean beech forests 91Y0 Dacian oak & hornbeam forests 91Z0 Moesian silver lime woods 91AA Eastern white oak woods 91BA Moesian silver fir forests 91CA Rhodopide and Balkan Range Scots pine forests Mediterranean and Macaronesian forests 9210 Apeninne beech forests with Taxus and Ilex 9220 Apennine beech forests with Abies alba and beech forests with Abies nebrodensis 9230 Galicio-Portuguese oak woods with Quercus robur and Quercus pyrenaica 9240 Quercus faginea and Quercus canariensis Iberian woods 9250 Quercus trojana woods 9260 Castanea sativa woods 9270 Hellenic beech forests with Abies borisii-regis 9280 Quercus frainetto woods 9290 Cupressus forests (Acero-Cupression) 9310 Aegean Quercus brachyphylla woods 9320 Olea and Ceratonia forests 9330 Quercus suber forests 9340 Quercus ilex and Quercus rotundifolia forests 9350 Quercus macrolepis forests 9360 Macaronesian laurel forests (Laurus, Ocotea) 9380 Forests of Ilex aquifolium 9390 Scrub and low forest vegetation with Quercus alnifolia 93A0 Woodlands with Quercus infectoria (Anagyro foetidae-Quercetum infectoriae) Mountainous coniferous forests 9410 Acidophilous Picea forests of the montane to alpine levels (Vaccinio-Piceetea) 9420 Alpine Larix decidua and/or Pinus cembra forests 9430 Subalpine and montane Pinus uncinata forests 9510 Southern Apennine Abies alba forests 9520 Abies pinsapo forests 9530 (Sub-) Mediterranean pine forests with endemic black pines 9540 Mediterranean pine forests with endemic Mesogean pines 9550 Canarian endemic pine forests 9560 Endemic forests with Juniperus spp

9570 Tetraclinis articulata forests 9580 Mediterranean Taxus baccata woods 9590 Cedrus brevifolia forests (Cedrosetum brevifoliae) 95A0 High oro-Mediterranean pine forests 5. GROUP 5: Steppe, heath and scrub habitats Habitat type code as referred to in Annex I to Directive 92/43/EEC Habitat type name as referred to in Annex I to Directive 92/43/EEC Salt and gypsum steppes 1430 Halo-nitrophilous scrubs (Pegano-Salsoletea) 1510 Mediterranean salt steppes (Limonietalia) 1520 Iberian gypsum vegetation (Gypsophiletalia) Temperate heath and scrub 4050 Endemic macaronesian heaths 4060 Alpine and Boreal heaths 4070 Bushes with Pinus mugo and Rhododendron hirsutum (Mugo-Rhododendretum hirsuti) 4080 Sub-Arctic Salix spp. scrub 40A0 Subcontinental peri-Pannonic scrub 40B0 Rhodope Potentilla fruticosa thickets 40C0 Ponto-Sarmatic deciduous thickets Sclerophyllous scrub (matorral) 5110 Stable xerothermophilous formations with Buxus sempervirens on rock slopes (Berberidion p.p.) 5120 Mountain Cytisus purgans formations 5140 Cistus palhinhae formations on maritime wet heaths 5210 Arborescent matorral with Juniperus spp

5220 Arborescent matorral with Zyziphus 5230 Arborescent matorral with Laurus nobilis 5310 Laurus nobilis thickets 5320 Low formations of Euphorbia close to cliffs 5330 Thermo-Mediterranean and pre-desert scrub 5410 West Mediterranean clifftop phryganas (Astragalo-Plantaginetum subulatae) 5420 Sarcopoterium spinosum phryganas 5430 Endemic phryganas of the Euphorbio-Verbascion 6. GROUP 6: Rocky and dune habitats Habitat type code as referred to in Annex I to Directive 92/43/EEC Habitat type name as referred to in Annex I to Directive 92/43/EEC Sea cliffs, beaches, and islets 1210 Annual vegetation of drift lines 1220 Perennial vegetation of stony banks 1230 Vegetated sea cliffs of the Atlantic and Baltic Coasts 1240 Vegetated sea cliffs of the Mediterranean coasts with endemic Limonium spp

1250 Vegetated sea cliffs with endemic flora of the Macaronesian coasts 1610 Baltic esker islands with sandy, rocky and shingle beach vegetation and sublittoral vegetation 1620 Boreal Baltic islets and small islands 1640 Boreal Baltic sandy beaches with perennial vegetation Coastal and inland dunes 2110 Embryonic shifting dunes 2120 Shifting dunes along the shoreline with Ammophila arenaria (‘white dunes’) 2130 Fixed coastal dunes with herbaceous vegetation (‘grey dunes’) 2140 Decalcified fixed dunes with Empetrum nigrum 2150 Atlantic decalcified fixed dunes (Calluno-Ulicetea) 2160 Dunes with Hippophaë rhamnoides 2170 Dunes with Salix repens ssp. argentea (Salicion arenariae) 2180 Wooded dunes of the Atlantic, Continental and Boreal region 2190 Humid dune slacks 2210 Crucianellion maritimae fixed beach dunes 2220 Dunes with Euphorbia terracina 2230 Malcolmietalia dune grasslands 2240 Brachypodietalia dune grasslands with annuals 2250 Coastal dunes with Juniperus spp